The black bellied seed cracker, Pyrenestes, is a West African finch. Within the same geographic region, two subspecies of the finch are found. One subspecies has a large beak, which is efficient at cracking the hard seeds of the sedge, Scleria verrucosa. The other subspecies has a small beak, which is more efficient at eating the soft seeds of the sedge, Scleria goossensii. What type of selection occurred to produce this situation